The family Euphorbiaceae Jussieu

Remark The words or terms in red (actually dark orange) in the text are defined in a glossary.


jun_07_02.gthmb can't be loaded. The Euphorbiaceae is a large family of plants, perhaps the sixth largest family, with about 240 genera and around 6 000 species. There is a large variation within the genus; most species are herbs, but some especially in the tropics, are also shrubs or trees, and some are lianas. Their flowers are usually actinomorphic (radially symmetrical); they are aggregated in terminal or axillary inflorescences that are always cymose. The flowers are unisexual; the female flowers have a superior ovary; the male flowers have only one stamen. The leaves are usually alternate, seldom opposite, and usually with stipules, that may be reduced to hairs, glands, or spines. The fruit is usually a capsular schizocarp. The family is characterized by the frequent occurrence of milky sap.

There are about 60 genera of Euphorbiaceae in the USA but only two genera are found in Québec:

Name

The family name is from the Euphorbia, the type genus. As for Euphorbia it is named for Euphorbus, the corpulent Greek juba.jpg can't be loaded. physician of Juba II (about 50 BC to bout 20 AD, picture at right), a Berber King of Numidia (present day Tunisia and eastern Algeria) and then later moved to Mauretania (present day western Algeria, and northern Morocco). Juba was educated in Rome, raised by Julius Caesar, and he took part in Caesar's triumphal procession. He married the daughter of the Roman triumvir Mark Antony and of the Queen Cleopatra VII of Egypt (the Cleopatra!). He was interested in botany and according to Pliny the Elder (AD 23 - August 24, pline.jpg can't be loaded. AD 79, picture at left), had written about an African cactus-like plant he had found or which he knew about from the slopes of Mount Atlas which was used as a powerful laxative. Galen mentions a short treatise written by the king on the virtues of the plant. That plant may have been Euphorbia resinifera or Euphorbia regisjubae (King Juba's Euphorbia) or Euphorbia officinarum and like all Euphorbias had a latexy exudate. Euphorbus had a brother, Antonius Musa, who was the physician to Augustus Caesar in Rome. When Juba heard that Caesar had honored his physician with a statue, he decided to honor his own physician by naming the plant he had written about after him.

In his Naturalis Historia, [XXV,38 (xxxviii, 77,78,79)] Pliny the Elder writes:


	Invenit et patrum nostrorum aetate rex Iuba quam appellavit Euphorbeam
	medici sui nomine. frater is fuit Musae, a quo divum Augustum conservatum
	indicavimus. iidem fratres instituere a balineis frigida multa corpora
	adstringere; antea non erat mos nisi calida tantum lavari, sicut apud
	Homerum etiam invenimus.

	Sed Iubae volumen quoque extat de ea herba et clarum praeconium. invenit
	eam in monte Atlante, specie thyrsi, foliis acanthinis. vis tanta est, ut e
	longinquo sucus excipiatur incisa conto; subitur excipulis ventriculo
	haedino. umor lactis videtur defluens; siccatus cum coiit, turis effigiem
	habet. qui colligunt, clarius vident. contra serpentes medetur quacumque
	parte percussa vertice inciso et medicamento addito ibi.

	Gætuli, qui legunt, taedio lacte adum non est, fastidiendum odorem habet.
	multum infra hunc sucum est qui in Gallia fit ex herba chamelaea, granum
	cocci ferente. fractus hammoniaco similis est, etiam levi gustu os accensum
	diu detinens et magis ex vero, donec fauces quoque siccet. 
that can be translated, literarily, (but not literaly):

	In the time, too, of our fathers, King Juba discovered a plant, to which he
	gave the name of euphorbia, in honour of his physician, Euphorbus, the 
	brother of the same Musa, whom We have mentioned as having saved the life
	of the late Emperor Augustus. It was these brothers who introduced the
	practice of douching the body with large quantities of cold water,
	immediately after the bath, for the purpose of bracing the system:
	whereas in former times, as we find stated in the works of Homer even, it
	was the practice to wash the body with warm water only.

	With reference to euphorbia, there is a treatise still in existence, written
	upon it by King Juba, in which he highly extols its merits: he discovered it
	growing upon Mount Atlas, and describes it as resembling a thyrsus in
	appearance, and bearing leaves like those of the acanthus. The properties of
	this plant are so remarkably powerful, that the persons engaged in
	collecting the juices of it are obliged to stand at a considerable distance.
	The incisions are made with a long pole shod with iron, the juice flowing
	into receivers of kid-leather placed beneath. The juice has all the 
	appearance of milk, as it exudes, but when it has coagulated and dried, it
	assumes the form and consistency of frankincense. The persons engaged in 
	collecting it, find their sight improved thereby. This juice is an excellent
	remedy for the stings of serpents: in whatever part of the body the wound
	may have been inflicted, the practice is to make an incision in the crown of
	the head, and there introduce the medicament.

	The Gætuli who collect it, are in the habit of adulterating it with warm
	milk; a fraud, however, easily to be detected by the agency of fire, that 
	which is not genuine emitting a most disgusting smell. Much inferior to this
	is the juice extracted, in Gaul, from the chamelæa, a plant which bears the 
	grain of Cnidos. When broken asunder, it resembles hammoniacum in
	appearance; and however slightly tasted, it leaves a burning sensation in 
	the mouth, which lasts a considerable time, and increases every now and
	then, until, in fact, it has quite parched the fauces.
That also illustrates how Latin could say much in few words!

In classical Greek ευφορβοσ (euphorbos) means well fed and some sources suggest that Juba was amused by the play upon words and chose his physician's name for the plant because of its succulent nature and because of Euphorbus corpulent physique.

Distribution

map_world.jpg can't be loaded. The family is cosmopolitan but occurs mainly in the tropics, with the majority of the species in the Indo-Malayan region and tropical America a good second. There is a large variety in tropical Africa, but it is not as abundant or varied as in these two other tropical regions. However, the Euphorbia genus also has many species in non-tropical areas such as the Mediterranean, the Middle East, South Africa and southern USA. The map shows the world distribution for the family.

Notes

The Euphorbiaceae family contains a large variety of phytotoxins, mainly diterpene esters, alkaloids and glycosides.

A number of plants of the Euphorbiaceae family are of considerable economic importance:

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